ARTICLES ON HUMAN ANATOMY, BONES OF THE UPPERLIMB (PHALANGES)
INTRODUCTION
The phalanges are digital bones in the hands and feet of most vertebrates. In primates, the thumbs and big toes have two phalanges while the other digits have three phalanges. The phalanges are classed as long bones.
The type of bones which form the skeleton of digits i.e. the fingers
and toes of the body are primarily known as phalanges bones. The human
body has a total of fifty-six phalanges, with three phalanges for each
finger and toes except for two phalanges per thumb and large toes in
feet. The phalanges are normally linked with each other at hinge-like
inter-phalangeal joints which is for extension purpose. In many cases,
the phalange bones are fused with each other.
Structure
The phalanges are the bones that make up the fingers of the hand and
the toes of the foot. There are 56 phalanges in the human body, with
fourteen on each hand and foot. Three phalanges are present on each
finger and toe, with the exception of the thumb and large toe, which possess only two. The middle and far phalanges of the fourth and fifth toes are often fused together (symphalangism). The phalanges of the hand
are commonly known as the finger bones. The phalanges of the foot
differ from the hand in that they are often shorter and more compressed,
especially in the proximal phalanges, those closest to the body.
A phalanx is named according to whether is are proximal, middle, or distal
and its associated finger or toe. The proximal phalanges are those that
are closest to the hand or foot. In the hand, the prominent, knobby
ends of the phalanges are known as knuckles. The proximal phalanges join with the metacarpals of the hand or metatarsals of the foot at the metacarpophalangeal joint or metatarsophalangeal joint.
The intermediate phalanx is not only intermediate in location, but
usually also in size. The thumb and large toe do not possess a middle
phalanx. The distal phalanges are the bones at the tips of the fingers
or toes. The proximal, intermediate, and distal phalanges articulate
with one another through interphalangeal articulations.
Bone anatomy
Each phalanx consists of a central part, called the body, and two extremities.
- The body is flat on either side, concave on the palmar surface, and convex on the dorsal surface. Its sides are marked with rough areas giving attachment to fibrous sheaths of flexor tendons. It tapers from above downwards.
- The proximal extremities of the bones of the first row present oval, concave articular surfaces, broader from side to side than from front to back. The proximal extremity of each of the bones of the second and third rows presents a double concavity separated by a median ridge.
- The distal extremities are smaller than the proximal, and each ends in two condyles (knuckles) separated by a shallow groove; the articular surface extends farther on the palmar than on the dorsal surface, a condition best marked in the bones of the first row.
In the foot, the proximal phalanges have a body that is compressed
from side to side, convex above, and concave below. The base is concave,
and the head presents a trochlear surface for articulation with the
second phalanx. The middle are remarkably small and short, but rather
broader than the proximal. The distal phalanges, as compared with the
distal phalanges of the finger, are smaller and are flattened from above
downward; each presents a broad base for articulation with the
corresponding bone of the second row, and an expanded distal extremity
for the support of the nail and end of the toe.
Distal phalanx
In
the hand, the distal phalanges are flat on their palmar surface, small,
and with a roughened, elevated surface of horseshoe form on the palmar
surface, supporting the finger pulp. The flat, wide expansions found at the tips of the distal phalanges are
called apical tufts. They support the fingertip pads and nails. The phalanx of the thumb has a pronounced insertion for the flexor pollicis longus
(asymmetric towards the radial side), an ungual fossa, and a pair of
unequal ungual spines (the ulnar being more prominent). This asymmetry
is necessary to ensure that the thumb pulp is always facing the pulps of
the other digits, an osteological configuration which provides the
maximum contact surface with held objects.
In the foot, the distal phalanges are flat on their dorsal surface.
It is largest proximally and tapers to the distal end. The proximal part
of the phalnx presents a broad base for articulation with the middle
phalanx, and an expanded distal extremity for the support of the nail
and end of the toe. The phalanx ends in a crescent-shaped rough cap of bone epiphysis —
the apical tuft (or ungual tuberosity/process) which covers a larger
portion of the phalanx on the volar side than on the dorsal side. Two
lateral ungual spines project proximally from the apical tuft. Near the
base of the shaft are two lateral tubercles. Between these a V-shaped
ridge extending proximally serves for the insertion of the flexor pollicis longus. Another ridge at the base serves for the insertion of the extensor aponeurosis. The flexor insertion is sided by two fossae — the ungual fossa distally and the proximopalmar fossa proximally.
Development
The
number of phalanges in animals is often expressed as a "phalangeal
formula" that indicates the numbers of phalanges in digits, beginning
from the innermost medial or proximal. For example, humans have a
2-3-3-3-3 formula for the hand, meaning that the thumb has two phalanges, whilst the other fingers each have three.
In the distal phalanges of the hand the centres for the bodies appear
at the distal extremities of the phalanges, instead of at the middle of
the bodies, as in the other phalanges. Moreover, of all the bones of
the hand, the distal phalanges are the first to ossify.
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